Though they were likely naturally rare, scientists believe that destructive fishing practices have reduced the numbers of Atlanticgoliath groupers by at least 80% and that the species is now critically endangered. These fish utilize the same, few locations and same, few days for spawning every year, so their presence is quite predictable.
Furthermore, a total lack of fear of people makes them an easy target for spear fishers. Finally, the Atlanticgoliathgrouper ’s large size, slow growth, and ease of capture all contribute to slow its recovery, even where laws have been put in place to give it some or complete legal protection from fishing (e.g., in the USA and Brazil).
Scientists only recently divided the species into two, based on their slightly different genetic makeup. The two species are similar in both appearance and behavior, but little is known about the population trends or conservation status of the Pacific goliathgrouper.
The once common Nassau grouper (Epimetheus stratus) and goliathgrouper (E. Tamara) have been so depleted that they are under complete protection from the South Atlantic, Gulf of Mexico, and Caribbean. From 1997-2005, our researchers collaborated with Florida State University's Institute for Fishery Resource Ecology (Dr. Chris Koenig and Dr. Felicia Coleman) to monitor the status and recovery of goliathgrouper.
This goliathgrouper research program investigated juvenile and adult Jewish abundance, distribution and migration patterns; their age and growth; and their habitat utilization. With the help of Don Maria we have tagged over 1,000 adult Jewish and have observed aggregations of goliathgrouper in both the Gulf of Mexico and more recently, the South Atlantic.
Posters created by the Center of Marine Conservation help disseminate information about our project and its requirements, highlighting our tagging study and the morphology of goliathgrouper. Given that these groupers were afforded protected status, researchers worked to utilize and develop novel non-lethal techniques to procure and analyze biological samples for life history information.
These casualties, resulting from red tide, gave our biologists a unique opportunity to collect a multitude of biological samples, without having to sacrifice healthy animals. From these decomposing carcasses, biologists were able to record length for use in an age/length relationship, and were able to extract monoliths and remove dorsal spines and rays for comparison of hard parts in age and growth analysis.
Tissue samples were also removed and sent to the Florida Marine Research Institute, so they could evaluate the level of red tide toxin. The sampling trip gave these biologists an opportunity to educate the curious beach goers about red tide and goliathgrouper (a few of which had been misidentified as baby manatees).
Attempts to evaluate the data needed to assess the status of these depleted stocks and develop rebuilding plans present unique challenges. In 2010, the Florida Fish and Wildlife Conservation Commission and NOAA Fisheries convened a benchmark goliathgrouper assessment for the continental U.S. population.
This project would not have been possible without ongoing collaboration with researchers from Florida State University, Everglades National Park, and the recreational fishing and SCUBA diving communities. The AtlanticGoliathGrouper is an endangered species of grouper found in the Florida Keys, the Bahamas, most of the Caribbean, and almost all the Brazilian coast.
Lurking in the deepest recesses of inshore waters is one of the most powerful and challenging species sought by anglers. With nearly no natural predators once adulthood is reached and with a history of even stalking human swimmers on occasion, this monster of the deep is truly a worthy adversary for any fisherman.
Goliath Groupers feed primarily on crustaceans such as spiny lobsters, shrimp, and crabs, as well as stingrays, octopus, and even young sea turtles, all of which it can easily catch and devour with its three to five rows of teeth. Popular locations to fish for GoliathGrouper include bridges and structure when angling inshore, and sunken wrecks and reefs offshore.
The easily approachable nature of the grouper makes it a great fish for spear fishermen, though this has reduced its population in areas accessible to divers. It is a full contact sport that can leave even the strongest and fittest of anglers exhausted only minutes into the fight.
However, if you’re truly dedicated to muscling one of these mammoths to the boat, and you can endure the long and demanding battle of catching one, you’ll be rewarded with a prize like no other. Be sure to take lots of pictures, because you’ll definitely want proof to back up your story when regaling your jealous friends with your tale of triumph.
Groupers are generally a friendly species and can be found patrolling artificial and coral reefs alike, primarily in shallow tropical waters. Goliath groupers navigate to an annual spawning for breeding, the season and location varies depending on the population.
In Florida, hatchlings join their brethren in safe spaces near coastal mangrove estuaries and spend their first six years of life dining exclusively on fish, crabs, and shrimp before heading out to open waters. The Goliath grouper grows slowly, attaining maturity around age 20-25, which is why it is important to manage fishing of the species; they need the chance to reach adulthood to reproduce in order to create a sustainable fishery.
The Goliath grouper is a key species in Florida waters because their presence is an indicator of health for local coral reefs. This particular species feeds by swallowing their prey whole, creating negative pressure that quickly them to bring in whole invertebrates, fish, and even smaller sharks.
Many grouper, manatees, and turtles were found washed ashore on Southwest Florida beaches during the red tides in 2003 and 2005. The good news is that as of 2006 the Goliath grouper ’s population had improved and was considered to be on a recovery trajectory due to the careful protection by NOAA Fisheries.
It possesses a robust and elongated body, with a wide head in comparison to its small eyes. What’s more, the base of the dorsal fin stands out as being covered with scales and thick skin.
Its yellowish, grayish or olive-toned coloring with small spots help the Atlanticgoliathgrouper blend into its environment. The majority of these gigantic fish live in deep waters, near rocky areas with coral and mud.
Their geographic location is, for the most part, the American coast from Florida down to southern Brazil. However, a number of these fish also live along the Pacific coast from California down to Peru.
It’s also worth pointing out that there are also Atlanticgoliath groupers living along the African coast from Senegal to the Congo. The enormous size of this fish, along with its great gastronomic value, make it a much sought-after catch for fishermen.
However, its slow growth and low reproduction rate make it one of the most susceptible species to extinction. The Atlanticgoliathgrouper, whose scientific name is Epimetheus Tamara, is a grouper species characterized by its large dimensions.
Historically, its size has made it popular among fishers and merchants to the point that it became the object of recreational competitions. Furthermore, its meat stands out for its nutritional value and renowned taste, similar to that of the common grouper.
The Atlanticgoliathgrouper or Tamara (Epimetheus Tamara), also known as the Jewish, is a large saltwater fish of the grouper family found primarily in shallow tropical waters among coral and artificial reefs at depths from 5 to 50 m (16 to 164 ft). Its range includes the Florida Keys in the US, the Bahamas, most of the Caribbean and most of the Brazilian coast.
On some occasions, it is caught off the coasts of the US states of New England off Maine and Massachusetts. In the eastern Atlantic Ocean, it occurs from the Congo to Senegal.
Scientific classification Kingdom: Animalia Phylum: Chordata Class: Actinopterygii Order: Performed Family: Serranidae Subfamily: Epinephrine Genus: Epimetheus Species: Binomial name Epimetheus Tamara Synonyms Promiscuous Tamara (Lichtenstein, 1822) Serra nus Tamara Lichtenstein, 1822 Serra nus Menelik Valentines, 1828 Serra nus gales J.P. Müller & Trochee, 1848 Serra nus guava Play, 1860 Promiscuous one Ehrenberg, 1915 Promiscuous ditto Roux & Collision, 1954 Young Atlanticgoliath groupers may live in brackish estuaries, oyster beds, canals, and mangrove swamps, which is unusual behavior among groupers.
They may reach extremely large sizes, growing to lengths up to 2.5 m (8.2 ft) and can weigh as much as 360 kg (790 lb). The world record for a hook-and-line-captured specimen is 308.44 kg (680.0 lb), caught off Fernanda Beach, Florida, in 1961.
Considered of fine food quality, Atlanticgoliathgrouper were a highly sought-after quarry for fishermen. It is a relatively easy prey for spear fishermen because of the grouper's inquisitive and generally fearless nature.
They also tend to spawn in large aggregations, returning annually to the same locations. This makes them particularly vulnerable to mass harvesting while breeding.
Until a harvest ban was placed on the species, its population was in rapid decline. The fish is recognized as “vulnerable” globally and “endangered” in the Gulf of Mexico.
The species' population has been recovering since the ban; with the fish's slow growth rate, however, some time will be needed for populations to return to their previous levels. Goliath groupers are believed to be protogynous hermaphrodites, which refer to organisms that are born female and at some point in their lifespans change sex to male.
Most grouper follow this pattern, but this has not yet been verified for the Goliath. Males can be sexually mature at about 115 centimeters (45 in), and ages 4–6 years.
Tidal pools act as nurseries for juvenile E. Tamara. In tidal pools juvenile E.Tamara are able to utilize rocky crevices for shelter.
Besides shelter, tidal pools provide E. Tamara with plenty of prey such as lobster and porcelain crab. It may have referred to the fish's status as inferior leading it to be declared only suitable for Jews, or the flesh having a “clean” taste comparable to kosher food ; it has also been suggested that this name is simply a corruption of jaw fish or the Italian word for “bottom fish”, Giuseppe.
In 2001, the American Fisheries Society stopped using the term because of complaints that it was culturally insensitive. IUCN Red List of Threatened Species.
^ Lovato, Cleo nice Maria Cardozo; Soars, Bruno Clears; Begot, Tiago Octavio Buffalo; Montage, Luciano Coach de Assis (January 2016). “Tidal pools as habitat for juveniles of the Goliath grouper Epimetheus Tamara (Lichtenstein 1822) in the Amazonian coastal zone, Brazil”.
Risky, Delaney C.; Bakenhaster, Micah D.; Adams, Douglas H. (2015). “ Pseudorhabdosynochus species (Monogenoidea, Diplectanidae) parasitizing groupers (Serranidae, Epinephrine, Epinephrine) in the western Atlantic Ocean and adjacent waters, with descriptions of 13 new species”.
Southern stingrays, a green moray eel, and two sea turtles also share this exciting exhibit. Sand tiger sharks are found in the shallow, coastal tropical waters of all oceans except the central and eastern Pacific.
Diet This nocturnal hunter feeds on bony fish, small sharks and rays, octopus, and large crustaceans. Unique Adaptations Sand tiger sharks are often found in groups of a few dozen, hovering in caves or near reefs or shipwrecks.
Sand tigers migrate, coming toward shore during the summer and moving southward or to deeper waters in the winter. Reproduction Female sand tiger sharks are viviparous (producing living young from eggs that hatch within the body).
Southern stingrays inhabit temperate waters of bays and estuaries from New Jersey to Brazil, as well as the Gulf of Mexico. Diet Rays feed on a wide variety of bottom organisms, such as crustaceans (shrimp and crab), mollusks (snails and shellfish), and worms.
Development is viviparous (pups hatch from their egg capsules while inside the mother’s uterus and are born soon after). Sea turtles are air-breathing reptiles found in the tropical and temperate waters of the world’s oceans.
In the early spring and summer, females may leave the water and return to their home beach to nest. Kemp’s Ridley sea turtles are carnivores, feeding on crabs, mollusks, jellyfish, mussels, and fish.
Unique Adaptations Streamlined bodies and flippers make these large animals powerful swimmers and divers-some species routinely diving to depths greater than 1,000 feet and staying underwater for several hours. Unlike their freshwater relatives, sea turtles have a special gland that rids their bodies of excess salts.
These events are called an “arrived” and take place on a small strip of beach at Ranch Nero, Mexico. Their populations have declined due to commercial harvest of turtle meat, eggs, skins for leather, and shells for ornaments and jewelry.
They die from ingesting marine debris, such as plastic bags, or get caught in nets as by catch and drown. Moray eels are found in tropical reefs and shallows from New Jersey to Brazil, including Bermuda and the Gulf of Mexico.
They feed on fish and crustaceans, especially crabs, and use their long, slender bodies to enter holes and crevices in search of hidden prey. Despite warning predators with a large open mouth and sharp teeth, morays are not aggressive and do not usually bite unless provoked.
Moray eels begin adult life as males, then later change to females (sex reversal). They frequently follow large animals like sea turtles, sharks, and rays to scavenge leftovers.
They live in the tropical and temperate waters of the Atlantic Ocean, along the eastern coast of the U.S. and south to Brazil and Uruguay. Jacks are typically light, usually silver with red pigmentation which disappears in dark ocean water.
Diet These strong-swimming carnivores rely on speed and strength to catch their prey, which includes small fish, cope pods, and other ocean animals. Unique Adaptations Jacks are often found swimming with sharks, but when roaming the open sea they school as a defense mechanism.
Jacks can also be seen hitching a free ride in the bow wake of their predatory “companions.” Juveniles hide among jellyfish, debris, plants, etc, and have a deeper and sleeker appearance then the adults. Atlanticgoliath groupers are found in tropical and subtropical waters of the Atlantic Ocean from North Carolina to Brazil, and throughout the Caribbean and Gulf of Mexico.
They prefer the sheltered habitats of coral reefs, and especially shady areas such as shipwrecks, rock ledges, and caves. Atlanticgoliathgrouper have a broad, flat head and mouth, and can grow up to 8 feet long and weigh over 700 pounds.
These hunters are not built for speed over long distances and prefer to ambush prey, rather than pursue it in open water. The diet of these large predators consists mainly of crab, lobster, fish, octopus, and young sea turtles.
Habits and Adaptations Atlanticgoliath groupers are often spotted or dark, allowing them to camouflage with their surroundings. Small groupers may be preyed upon by barracuda, king mackerel, moray eels, and sharks.
When threatened, an Atlanticgoliathgrouper will defend its territory with aggressive body language and a distinctly audible rumbling sound. These puffer fish primarily inhabit coral reefs and other warm shallow waters.
Usually brown, with some yellow on the underside, these stocky, slow swimming fishes have a large head and a “box-shaped” body. Diet Porcupine fish use their strong beaks to crush coral polyps, mollusks, crustaceans, crabs, and sea urchins.
Unique Adaptations Puffers hide in coral and as their name implies, can puff up two to three times their normal size by sucking air or water into a special chamber in their abdomen. Puffers employ a number of defenses to avoid getting eaten: they are covered with sharp spines, their internal organs contain an extremely toxic nerve poison, and when cornered they can bury themselves in the sand.
This important food fish can be found in warm coastal waters, from Massachusetts to Brazil, including Bermuda and the Gulf of Mexico. They have a triangular-shaped head and a notched tail, and are capable of inflicting injuries to unsuspecting fishermen with their well-developed teeth.
Tarpon are found in tropical and temperate waters along the eastern Atlantic coasts of North and South America. Unique Adaptations When swimming in oxygen-poor water, tarpons can gulp air from the surface using special lung-like bladders.
Initially the young head to shallow water where they become an intricate part of the plankton that drifts with ocean currents far from shore. Trigger fish usually spend a large portion of their lives near a coral reef-inhabiting areas in coastal waters from New York to Brazil.
Small eyes set high on a large, angular head and jaw give them a “bucktoothed” appearance. Diet Trigger fish have powerful jaws and teeth that allow them to easily crush hard-shelled prey like crustaceans, mollusks, coral, and sea urchins.
The Queen trigger fish will blow mouthfuls of water at sea urchins to flip them over and expose their softer underparts. Epimetheus Tamara may be a widespread, slow growing, and aggregating species that has undergone vital population reduction over the past three generations (40.5 years) calculable to be a minimum of 80th based on landings information and underwater visual censuses.
Despite clear and promising signs of recovery in us waters following the 1990 moratorium, the will increase in numbers noted area unit young and juvenile fish (the species takes five to six years to become sexually mature). The goliathgrouper occurs within the western Atlantic Ocean from Florida south to Brazil, including the Gulf of Mexico and also the Caribbean Sea.
This marine fish inhabits shallow, inshore waters with mud, rock or coral bottoms and is infrequently found below depths of 46 meters. It’s territorial close to areas of refuge like caves, wrecks, and ledges, displaying an open mouth and quivering body to intruders.
These teams occur at consistent sites like wrecks, rock ledges and isolated patch reefs during July, August and September. Studies have shown fish could move up to 62 miles (100 km) from inshore reefs to these spawning sites.
In southwest Florida, plausible entreaty behavior has been observed during the complete moons in August and September. Occurring in shallow, inshore waters to depths of 150 feet (46 m), the Epimetheus Tamara prefers areas of rock, coral, and mud bottoms.
It’s a classic apex predator, large, rare and solely some people occur on any given reef unit. As with other fish, the Atlanticgoliathgrouper is the host of several species of parasites, including the diplectanid monogenean Pseudorhabdosynochus Americans on its gills.
Calico crabs frame the bulk of their diet, with alternative invertebrate species and fish filling within the rest. Goliath grouper feed mostly on crustaceans (in particular spiny lobsters, shrimps and crabs), fishes (including stingrays and parrot fishes), octopus, and young ocean turtles.
Here I’ll be digging into the best species for aquariums, as well as diet, tank setup, tank mates, and breeding. Groupers are saltwater fish in the subfamily Epinephrine of the Serranidae family (in the order Performed).
The majority of grouper fish can be quite difficult to sex as there aren’t many external differences between males and females. The biggest male groupers usually control harems that consist of up to three to fifteen females.
If a smaller female grouper changes its sex before it can form a harem as a male, then its fitness will decrease. This reproductive strategy is related to groups spawning high amounts under habitat cover.
As I mentioned earlier, most groupers fish are not particularly exciting to look at, coming in a dull brown or green color. These more pretty specimens can come in bright red, blue, orange, green, and black, adorned with equally beautiful patterns.
The largest hook and line captured Goliath was found in Florida and weighed 309 kilograms. A lot of grouper species in the Epinephrine subfamily are popular in home aquariums, so these are the fishes I’ll be focusing on in this article.
If you’re thinking of keeping a grouper, then here are some varieties that are suitable for home aquarium life. Some grouper species can make good pets, while others are not suited to a life in captivity.
Species such as the giant grouper definitely don’t belong in aquariums as they can grow to a huge size. Compared to other saltwater fish species, groupers can grow pretty big and need a large tank.
While this shouldn’t be an issue for experienced aquariums who are knowledgeable about keeping saltwater fish, it could be problematic for those new to the fish keeping hobby. Additionally, groupers are marine fish, so their care is a little more complicated than freshwater or Goldwater species.
If you’ve never kept fish before, I’d advise setting up a tropical or Goldwater tank first to help you get to grips with things. You’ll need to make sure you have the time and money to keep groupers in both the immediate and the long run.
Most grouper species are robust and hardy fish, so they don’t typically have any problems during transport or added to an aquarium. Unlike some fish that are notoriously sensitive and difficult to keep, groupers are fairly tough.
Atlantic Goliath will also eat octopuses, baby sea turtles, barracudas, and even sharks. You need seafood; scallops, squid, shrimp, halibut, and snapper are some excellent food options to keep your groupers satisfied.
Bear in mind that these fish, like all predatory animals, require whole food items to thrive. Whole Meal You’ll need to make sure you offer a type of whole animal item at least once or twice a week to your grouper.
It’s also a good idea to soak your grouper ’s food in a fish-friendly vitamin supplement a few times a week, too. When it’s time to feed your groupers, you can simply throw their food into the tank and let them eat.
You also need to be aware of the ideal water parameters to provide optimal living conditions for your pet. Before you bring your grouper fish home, you’ll need to make sure your aquarium has everything they need to live comfortably.
Sandy substrates such as granite sand or crushed coral are best for grouper tanks. Gravel isn’t generally used in marine tanks as it lacks buffering capabilities.
If your filter lacks strength and power, it won’t be able to effectively clean your tank water. As a general rule of thumb, your filter should be able to clean at least four times the volume of your aquarium.
For example, if you have a 200-gallon grouper tank, then you’ll need a filter that has a flow rate of at least 800 gallons per hour (Mph). You’ll need to use an aquarium heater when keeping grouper species to ensure your tank water remains at a consistently warm temperature.
Unless you’re keeping corals and live plants, you don’t need an overly strong lamp. An aquarium light will help replicate a natural day and night cycle for your groupers.
If you want to keep corals and live plants, then you’ll require a more powerful light with the right color spectrum. Groupers need plenty of hiding spaces in their tank, so make sure their aquarium has a variety of caves and crevices they can retreat to.
Adding live plants to your grouper tank can also help them feel more comfortable and at ease. As groupers are carnivores, they won’t pay any attention to live plants, so you don’t need to worry about them destroying your aquas cape.
If you want to house a grouper in a reef tank, you’ll need to select your tank mates carefully to ensure they don’t become (expensive!) Many groupers can grow pretty big and long, so they require a decent-sized aquarium.
Before keeping a grouper, make sure your tank’s capacity is big enough for their adult size. Additionally, if you want to keep two or more groupers together, then you’ll need a larger aquarium than the minimum tank size for each species.
Like all species of fish, groupers require specific water parameters to survive in the home aquarium. As groupers are marine species of fish, they require salt water to survive.
As I mentioned earlier, groupers have pretty high bio-loads due to their big appetites and messy feeding habits. If your tank is small and under stocked, then you’ll need to perform larger and more frequent water changes.
You’ll need to make sure you have a fish tank heater to keep their environment at an appropriate temperature. Scallops, crayfish, halibut, and whole shrimp are some foods that you can feed them two times a day.
Health In addition to feeding your groupers, you’ll need to check their behavior every day and watch out for any signs of illness and disease. Checking your aquarium equipment for signs of damage and testing your tank water are other tasks to complete regularly.
If you want to keep a grouper in a community aquarium or reef tank, then you’ll need to choose their tank mates carefully. Some tank mates that work well in a grouper tank include trigger fish (with caution), large marine angelfish, tangs, lionfish, and rabbit fish.
This is an important step if you have another grouper or aggressive fish already present in the tank, like trigger fish and marine angelfish. Doing so removes established territories and puts all your fish in unfamiliar surroundings, which helps prevent any aggressive reactions towards your new grouper.
The Best Time To Transfer Additionally, introducing your grouper to your community tank in the evening when the lights have been off for a while can also reduce aggression. In fact, most groupers are kept alone in captivity as housing multiple ones together requires an incredibly big aquarium.
Groupers can develop illnesses or diseases that can affect their quality of life and lifespan. Groupers can develop illnesses or diseases that can affect their quality of life and lifespan.
Loss of appetite Reddening, torn, or fraying fins/tail Respiratory distress Red gills Antisocial behavior Stringy white feces Identifying possible causes behind your grouper ’s poor health is crucial for the right course of action and treatment.
This organism is naturally found in many tanks and is exceptionally resilient, making it hard to control. Fish suffering from marine velvet will often have symptoms like bleeding or inflammation of the gills, difficulty breathing, lethargy, rubbing against tank objects, and destruction of lung tissue.
Unfortunately, by the time these golden spots appear, the gills are usually irreversibly damaged and treatment is no longer effective. As marine velvet progresses, the gills and lung tissue will start to die off.
This can result in the fish gradually being unable to transport oxygen to the gill membranes, eventually leading to suffocation. It is important to note that copper is lethal to fish in high doses, so make sure you follow the manufacturer’s instructions and dosage guidelines carefully.
The exact cause of Hole in the Head Disease isn’t known, but it has been linked to poor water quality and nutritional deficiencies. Irregular water changes can also influence the development of Hole in the Head Disease.
You should offer your fish live and frozen foods with vitamin supplements and fresh vegetables. Improving the water quality in your aquarium is also mandatory for treating Hole in the Head Disease.
As it is an internal bacterial infection, many fish don’t show any external symptoms at first. At this point, the fish will usually display red streaks on the body (which often points to internal hemorrhaging), dark swollen lesions, cloudy eyes, red spots, lethargy, poor appetite, and respiratory distress.
It’s also important to note that fibrosis can be transmitted to humans by contacting infected fish, though this is quite rare. If you’re dealing with fish infected with this disease, make sure you avoid touching contaminated tank water and any cuts or lesions.
The breeding process for groupers varies between species, but many seem to spawn offshore on shelf and shelf-edge reefs. Their pelagic larvae stay in the open ocean for around 40 to 60 days, at which point they will reach inshore nursery grounds.
The larvae turn into small juveniles and stay in the inshore nursery grounds for a fairly long period. For example, male Atlantic Goliath are considered mature once they are over 7 years old and are at least 45.5 inches long.
Spawning for Atlantic Goliath tends to happen during July, August, and September. Goliath's prefer to spawn in isolated patch reefs, shipwrecks, and rock ledges.
After around 25 to 26 days after hatching, the pelagic larvae transform into benthic juveniles and are just 1-inch long. Fry have tiny yolk sacs when they are newly hatched, so they will begin to eat almost instantly.
These fish are slow growers and require at least 7 months to reach adult coloration. As I mentioned earlier, most groupers are monastic protogynous hermaphrodites, which means they have the ability to change sex.
Many groupers of species can be bred in captivity, though not a lot of information is available on how to breed them in home aquariums. Groupers are incredibly hard to sex as there doesn’t appear to be any external differences between males and females.
If you want a unique and hardy centerpiece fish for your marine tank, then groupers are well worth considering. These fish have huge appetites and heavy bio loads, so good filtration is key.
They are carnivores, so feed them a good variety of meaty foods such as scallops, whole shrimp, mussels, and squid every day. As long as you have the time, space, and experience to keep a grouper fish, I think they’re a great species to add to a saltwater tank.